This is the typical posture of a territorial call. ©MRR 2017
The white-tailed prairie dog will sometimes throw its head back in a territorial call. ©John Hoogland 2006
And other times the white-tailed prairie dog will not throw its head back. ©John Hoogland 2006
It's approaching evening and 45, the dominant male in his clan, lets out a territorial call. ©MRR 2017
There are a few postures given for territorial calls, but a forward head and half-open mouth in an O is the tell-tale visual sign. ©MRR 2017
45 lets out a passionate territorial call mid-run. ©MRR 2017
Though not as often as males, females also let out territorial calls throughout the day. ©MRR 2017
A white-tailed prairie dog territorial call. ©John Hoogland 2006
The head-back posture can be extreme in the white-tailed species. ©John Hoogland 2007
R21 stops mid-fight to let out a territorial call. A clear scar on his left cheek was acquired from one such fight with another male. ©MRR 2017
0 (zero), the dominant male in his clan, spends all day during the mating season running around and establishing his territory. ©MRR 2017
The classic O mouth shape can be seen here as 0 lets out a territorial call mid-run. ©MRR 2017
A classic territorial call posture. ©MRR 2017
Snow-covered mornings see prairie dogs waking up late and slow, but the dominant male always comes out early and lets out a territorial call. ©MRR 2017
During the mating season, males will visit the burrows in their territory looking for females and establishing their dominance. ©MRR 2017
Another white-tailed prairie dog giving a territorial call without the head thrown back. ©John Hoogland 2010
R21 did a lot of traveling as he attempted to secure his own territory, letting out frequent territorial calls. ©MRR 2017
Another head-back territorial call by a white-tailed prairie dog. ©John Hoogland 2010
This rare image shows a Gunnison's prairie dog throwing his head back during a territorial call. The head throw is not known within this species, but this particular individual (adult male 5) did it often. ©MRR 2017
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